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Tarkhnishvili, David (1996) The distribution and ecology of the amphibians of Georgia and the Caucasus: a biogeographical analysis. Zeitschrift für Feldherpetologie, 3. С. 167-196. ISSN 0946-7998
Tarkhnishvili, David და Gokhelashvili, Ramaz (1996) A contribution to the ecological genetics of frogs: age structure and frequency of striped specimens in some Caucasian populations of the Rana macrocnemis complex. Alytes, 14 (1). С. 27-41. ISSN 0753-4973