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Tarkhnishvili, David (1993) Anurans of Borjomi Canyon: clutch parameters and guild structure. Alytes, 11 (4). С. 140-154. ISSN 0753-4973
Tarkhnishvili, David და Serbinova, Irina (1993) The Ecology of the Caucasian Salamander (Mertensiella caucasica Waga) in a Local Population. Asiatic Herpetological Research, 5. С. 147-145. ISSN 1051-3825