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Tarkhnishvili, David and Thorpe, Roger S and Arntzen, J W (2000) Pre-Pleistocene Refugia and Differentiation between Populations of the Caucasian Salamander. Molecular Phylogenetics and Evolution, 14 (3). pp. 414-422.
Tarkhnishvili, David and Arntzen, J W and Thorpe, Roger S (1999) Morphological variation in brown frogs from the Caucasus and the taxonomy of the Rana macrocnemis group. Herpetologica, 55 (3). pp. 406-417. ISSN 0018-0831
Arntzen, J W and Papenfuss, Theodore and Kuzmin, Sergius and Tarkhnishvili, David and Ishchenko, Vladimir and Tuniyev, Boris and Sparreboom, Max and Rastegar-Pouyani, Nasrullah and Ugurtas, Ismail and Anderson, Steven and Babik, Wiesiek and Miaud, Claude and Crnobrnja Isailovic, Jelka (2009) Triturus karelinii. IUCN, IUCN Red List of Threatened Species.