Multivariate analysis of body proportions, skin texture, and coloration characteristics of brown frogs (Rana macrocnemis and R. camerani) from 14 Georgian populations confirms the existence of two, geographically distinct groups of populations. Frogs in the first group (R. macronemis) are characterized by a smooth skin, a pale spotted pattern, absence of a mid-dorsal stripe, and a small inner metatarsal tubercle. Frogs in the second group (known as R. camerani) possess a rugose skin, a conspicuous pattern of dorsal spots, a mid-dorsal stripe with contrasting...